Invited book chapters

McGregor, P.K. and A.G. Horn. 2014. Communication Networks. In J. Krause, D. Croft, and D. James (eds.) Animal Social Networks: Perspectives and Challenges. Oxford University Press.

McGregor, P.K., A.G. Horn, M.L. Leonard, and F. Thomsen. 2013. Anthropogenic noise and conservation. In H. Brumm (ed.) Animal Communication and Noise. Springer Verlag, Berlin.

Horn, A.G. and P.K. McGregor. 2013. Information and influence in animal communication networks. In U. Stegmann (ed.) Animal Communication Theory. Cambridge University Press.

Horn, A.G. and M.L. Leonard. 2010. Parent-offspring signaling. In M. Breed and J. Moore (eds.), Encyclopedia of Animal Behavior. Elsevier Inc., Philadelphia.

Horn, A.G. and M.L. Leonard. 2005. Nestling begging as a communication network. In P.K. McGregor (ed.), Communication Networks and Animal Communication. Cornell Univ. Press, Ithaca, NY.

Horn, A.G. and M.L Leonard. 2002. Efficacy and the design of begging signals.   In J. Wright and M.L. Leonard (eds.), The Evolution of Begging.  Kluyver Academic Press, Dordrecht. Pp. 127-141.

Horn, A.G.  1997.  Speech acts and animal signals.  In D.W. Owings, M.D. Beecher, and N. Thompson (eds.), Perspectives in Ethology, Vol. 12, pp. 347-358.

Horn, A.G. and J.B. Falls.  1996.  Categorization and the design of signals.  In D.E. Kroodsma and T.E. Miller (eds.), Ecology and Evolution of Acoustic Communication in Birds, pp. 121-135.  Cornell Univ. Press, Ithaca, NY.

Staicer, C., D. Spector, and A.G. Horn.  1996.  Patterns of diel singing behavior in birds.  In D.E. Kroodsma and T.E. Miller (eds.), Ecology and Evolution of Acoustic Communication in Birds, pp. 426-453.  Cornell Univ. Press, Ithaca, NY.

Horn, A.G.  1992.  Field methods for studying the perception of song types by birds.  In P.K. McGregor (ed.), Playback experiments: issues and prospects, pp. 191-200.  Plenum, New York.

P.K. McGregor, C.K. Catchpole, T. Dabelsteen, J.B. Falls, C. Gerhardt, A.G. Horn, M. Lambrechts, D. Kroodsma, K. McComb, I. Pepperberg, L.M. Ratcliffe, W. Searcy, and D. Weary.  1992.  Issues in playback: the Thornbridge Hall consensus.  In P.K. McGregor (ed.), Playback experiments: issues and prospects.  Plenum, New York.

Refereed papers

Ronconi, R. A., J. R. Stephens, Z. J. Crysler, I. L. Pollet, D. T. Fife, A. G. Horn, and P. D. Taylor. 2015. Distribution, abundance and trends of gulls and terns breeding on Sable Island, Nova Scotia, Canada. Waterbirds [conditional acceptance]

Haff, T. M., A. G. Horn, M. L. Leonard, and R. D. Magrath. 2014. Conspicuous calling near cryptic nests: a review of hypotheses and a field study on white-browed scrubwrens. J. Avian Biol. DOI: 10.1111/jav.00622

McIntyre, E., M. L. Leonard, and A. G. Horn. 2014. Do nestling Tree Swallows (Tachycineta bicolor) respond to parental alarm calls? The Auk 131: 314-320.

Reers, H., M. Leonard, A. Horn, and A. Jacot. 2014. Parent-environmental interactions shape acoustic signatures in tree swallows: a cross-fostering experiment. Journal of Avian Biology 45: 123-130.

Winkler, D.W., K.M. Ringelman, P. O. Dunn, L. Whittingham, D. J. T. Hussell, R. G. Clark, R. D. Dawson, L.S. Johnson, A. Rose, S. H. Austin, W. D. Robinson, M. P. Lombardo, P. A. Thorpe, D. Shutler, R. J. Robertson, M. Stager, M. Leonard, A. G. Horn, J. Dickinson, V. Ferretti, V. Massoni, F. Bulit, J. C. Reboreda, M. Liljesthr�m, M. Quiroga, E. Rakhimberdiev, and D. R. Ardia. 2014. Latitudinal variation in clutch size-lay date regressions in Tachycineta swallows: effects of food supply or demography? Ecography 37:001-009.

McIntyre, E., M. L. Leonard, and A. G. Horn. 2013. Ambient noise and parental communication of predation risk in tree swallows, Tachycineta bicolor. Animal Behaviour 87: 85-89.

Laughlin, A.J., C. M. Taylor, D. W. Bradley, D. Leclair, R. G. Clark , R. D. Dawson , P. O. Dunn , A. Horn , M. Leonard, D. R. Sheldon, D. Shutler, L. A. Whittingham, D. W. Winkler, D. R. Norris. 2013.  Integrating information from geolocators, weather radar and citizen science to uncover a key stopover area for an aerial insectivore. The Auk 130: 23-239.

Fairhurst, L., A. G. Horn, and M. L. Leonard. 2013. Nest acoustics and begging call structure in nestling tree swallows. Animal Behaviour 85: 917-923.

Lucas, Z., A. Horn, and B. Freedman. 2012. Beached bird surveys on Sable Island, Nova Scotia, 1993 to 2009, show a decline in the incidence of oiling. Proceedings of the Nova Scotia Institute of Science 47: 91-129.

Zanette, L., M. Clinchy, M.L. Leonard, A.G. Horn, D.T. Haydon, and E. Hampson. 2012. Brood-parasite-induced female-biased mortality affects songbird demography: negative implications for conservation. Oikos 121: 1493-1500.

Farmer, R.G., M.L. Leonard, and A.G. Horn. 2012. Observer effects and avian call count survey quality: rare-species biases and overconfidence. The Auk 129: 76-86.

Leonard, M.L. and A.G. Horn. 2012. Ambient noise increases missed detections in nestling birds. Biology Letters 8: 530-532.

Shutler, D., D. Hussell, R. Norris, D. Winkler, R. Robertson, F. Bonier, W. Rendell, M. B�lisle, R. Clark, R. Dawson, N. Wheelwright, M. Lombardo, P. Thorpe, M. Truan, R. Walsh, M. Leonard, A. Horn, C. Vleck, D. Vleck, A. Rose, L. Whittingham, P. Dunn, K. Hobson, and M. Stanback. 2011. Spatiotemporal Patterns in Nest Box Occupancy by Tree Swallows Across North America. Avian Conservation and Ecology 7: 3. http://dx.doi.org/10.5751/ACE-00517-070103

Marques, P., M.L. Leonard, A.G. Horn and A. Contasti. 2011. How nestling tree swallows (Tachycineta bicolor) integrate their responses to hunger and signalling by nestmates. Ethology 117: 163-170.

Magrath, R.D, T.M. Haff, A.G. Horn and M.L. Leonard. 2010. Calling in the face of danger? How predation risk affects acoustic communication by parent birds and their offspring. Adv. Stud. Behav. 41:187-253.

Leonard, M.L., A.G. Horn and A. Dorland. 2009. Does begging call convergence increase feeding rates to nestling tree swallows? J. Avian Biol. 40: 243-247.

Horn, A.G. and M.L. Leonard. 2008. Acoustic interactions in broods of nestling birds (Tachycineta bicolor). J. Comp. Psychol. 122: 298-304.

Leonard, M. L. and A.G. Horn.  2006.  Age-related changes in signaling of need by nestling tree swallows (Tachycineta bicolor). Ethology 112: 1020-1026.


Leonard, M.L. and A.G. Horn. 2005. Ambient noise and the design of begging signals. Proc. Roy. Soc., Biol. Sci. 272: 651-656.

Leonard, M.L., A.G. Horn, and A. Mukhida. 2005. False alarms and begging in nestling birds. Anim. Behav. 69: 701-708.


Zink, R.M., J.D. Rising, S. Mockford, A.G. Horn, J.M. Wright, M. Leonard, and M.C. Westburg. 2005. Mitochondrial DNA variation, species limits, and rapid evolution of plumage coloration and size in the Savannah Sparrow. Condor 107: 21-28.


Shutler, D., M.L. Leonard, A.G. Horn, R. Shutler, and D.J.T. Hussell. 2004. Breeding between Tree Swallows from the same brood. J. Field Ornith. 74: 353-358.


Leonard, M. L., Horn, A. G. & Porter, J.  2003  Does begging affect growth in nestling tree swallows, Tachycineta bicolor? Behav. Ecol. Sociobiol. 54: 573-577.

Leonard, M.L., A.G. Horn, and E. Parks. 2003. The role of posturing and calling in the begging display of nestling birds. Behav. Ecol. Sociobiol. 54: 188-193.


Maurer, G., R. Magrath, M.L. Leonard, A.G. Horn, and C. Donnelly. 2003. Begging to differ: scrubwren nestlings beg to alarm calls and vocalize when parents are absent. Anim. Behav. 65: 1045-1055.


Leonard, M.L. and A.G. Horn. 2001. Acoustic signalling of hunger and thermal state by nestling tree swallows.  Anim. Behav. 61: 87-93.

Leonard, M.L. and A.G. Horn.  2001. Begging calls and parental feeding decisions in tree swallows (Tachycineta bicolor). Behav. Ecol. Sociobiol. 49: 170-175.

Leonard, M.L. and A.G. Horn.  2001. Begging in the absence of parents by nestling tree swallows. Behav. Ecol. 12: 501-505.


Leonard, M.L. and A.G. Horn. 2001.  Dynamics of calling by tree swallow (Tachycineta bicolor) nestmates. Behav. Ecol. Sociobiol. 50: 430-435.

Leonard, M.L., A. G. Horn, A. Gozna and S. Ramen. 2000.  Brood size and begging intensity in nestling tree swallows. Behav. Ecol. 196-201.

Leonard, M.L. and A.G. Horn.  1998.  Need and nestmates affect begging in tree swallows.  Behav. Ecol. 42: 431-436.

Leonard, M.L., A.G. Horn, C. Brown, and N. Fernandez.  1997.  Parent-offspring recognition in tree swallows.  Anim. Behav. 54: 1107-1116.

Horn, A.  1996.  Dawn song repertoires of tree swallows.  Can. J. Zool. 74: 1084-1091.

Leonard, M.L. and A.G. Horn.  1996.  Provisioning rules in tree swallows. Behav. Ecol. Sociobiol. 38: 341-347.

Leonard, M.L., J.L. Dickinson, A.G. Horn, and W.D. Koenig.  1996.  An experimental test of offspring recognition in western bluebirds.  Auk 112: 1062-1064.

Horn, A.G., M.L. Leonard, and D.M. Weary.  1995.  Oxygen consumption during crowing by roosters: talk is cheap.  Anim. Behav. 50: 1171-1175.

Leonard, M.L., A.G. Horn, and W.R. Fairfull. 1995. Causes and consequences of allopecking in white leghorn chickens.   Appl. Anim. Behav. Sci. 43:17-26.

Martin, K., Horn, A.G., and S.J. Hannon. 1995. The calls and associated behavior of breeding willow ptarmigan in Canada.  Wilson Bull. 107: 496-509.

Leonard, M.L. and A.G. Horn.  1995.    Crowing in relation to status in roosters.  Anim. Behav. 49: 1283-1290.


Leonard, M.L., K. Teather, A.G. Horn, W. Koenig, and J. Dickinson.  1994.  Sex-biassed provisioning in western bluebirds.  Beh. Ecol. 5:355-359.

Horn, A.G., T.E. Dickinson, and J.B. Falls.  1993.  Repertoires and male quality in western meadowlarks (Sturnella neglecta).  Can. J. Zool. 71: 1059-1061.

Martin, K. and A.G. Horn.  1993.  Clutch defense by male and female willow ptarmigan. Ornis Scand. 24: 261-266.

Horn, A.G., M.L. Leonard, L. Ratcliffe, R. Weisman, and S. Shackleton.  1992.  Variation in the whistled songs of black-capped chickadees.  Auk 109: 847-852.


McGregor, P.K. and A.G. Horn.  1992.  Strophe length and response to playback in great tits.  Anim. Behav. 43: 667-676.

Horn, A.G. and J.B. Falls.  1991.  Song switching in mate attraction and territory defense by western meadowlarks (Sturnella neglecta).  Ethology 87: 262-268.

Leonard, M.L., A.G. Horn, and L. Ratcliffe.  1991.  Parental aggression in black-capped chickadees.  Behav. Ecol. 2: 228-233.

Shackleton, S.A., L.M. Ratcliffe, A.G. Horn, and C.T. Naugler.  1991.  Song repertoires of Harris' sparrows (Zonotrichia querula).  Can. J. Zool. 69: 1867-1874.

Leonard, M.L., A.G. Horn, and S.F. Eden.  1989.  Does juvenile helping in moorhens enhance breeder reproductive success? A removal experiment.  Behav. Ecol. Sociobiol. 25: 357-361.

Horn, A.G. and J.B. Falls.  1988.  Structure of western meadowlark (Sturnella neglecta) song repertoires.  Can. J. Zool. 66: 284-288.

Horn, A.G. and J.B. Falls.  1988.  Repertoires and countersinging in western meadowlarks (Sturnella neglecta).  Ethology 77: 337-343.

Horn, A.G. and J.B. Falls.  1988.  Responses of western meadowlarks to song repetition and contrast.  Anim. Behav. 36: 291-293.

Horn, A.G.  1988.  Interspecific aggression in western meadowlarks (Sturnella neglecta): re-directed aggression?  J. Field Ornithol. 59: 224-226.

Horn, A.G. and J.B. Falls.  1986.  Western meadowlarks switch song types when matched by playback.  Anim. Behav. 34: 927-929.

Leonard, M.L., A.G. Horn, and S.F. Eden.  1988.  Parent-offspring aggression in moorhens.  Behav. Ecol. Sociobiol. 23: 265-270.

Eden, S.F., M.L. Leonard, and A.G. Horn.  1988.  Food provisioning lowers interclutch interval in moorhens Gallinula chloropus.  Ibis 131: 429-432.

Picman, J., M.L. Leonard, and A.G. Horn.  1988.  Anti-predation role of nest clumping in red-winged blackbirds.  Behav. Ecol. Sociobiol. 22: 9-15.

Weary, D.M., J.R. Krebs, R. Eddyshaw, P.K. McGregor, and A.G. Horn.  1988.  Decline in song output by great tits: exhaustion or motivation?  Anim. Behav. 36: 1242-1244.

Falls, J.B., A.G. Horn, and T.E. Dickinson.  1987.  How meadowlarks classify their songs: evidence from song matching.  Anim. Behav. 38: 579-585.

McGregor, P.K., A.G. Horn, and M.A. Todd.  1986.  Are familiar sounds ranged more accurately?  Perceptual and Motor Skills 61: 1082.

Invited book reviews

Horn, A.G. 2007. Deconstructing Animal Deceit (review of The Evolution of Animal Communication by W.A. Searcy and S. Nowicki). Ethology 113: 207-208.

Horn, A.G. 2006. [Multiple review of three books on bird song]. Bioacoustics.

Otter, K. A. and A. G. Horn. 2004. Animal Signals (by J. Maynard Smith and D. Harper). ISBE Newsletter 16 (2): 24-27.


Horn, A.G.  1998.   Music, Gestalt, and Computing: Studies in Systematic and Cognitive Musicology (ed. by M. Leman).  Bioacoustics 9: 239-241.

Horn, A.G.   1998.  Bird Song: Evolutionary Themes and Variations (by C.K. Catchpole and P.J.B. Slater).  Bioacoustics.

Popular articles

McLaren, I., and A.G. Horn. 2006. The Ipswich Sparrow: past, present, and future. Birding 38: 52-59.